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Three thousand years ago, warlord Yaotl enters a portal into a parallel universe and becomes immortal, but his four generals Aguila, Serpiente, Gato and Mono are turned to stone. The portal also releases 13 immortal monsters that destroy his army and his enemies.

In the present day, the Teenage Mutant Ninja Turtles have grown apart. After defeating their archenemy Shredder, Master Splinter has sent Leonardo to Central America for training, where he protects a village from bandits. Donatello works as an IT specialist, Michelangelo works as a birthday party entertainer known as "Cowabunga Carl" and Raphael secretly works at night as a vigilante known as "Nightwatcher". April O'Neil operates a unnamed company that locates and acquires relics for collectors, assisted by her boyfriend Casey Jones.

April travels to Central America for work and finds Leonardo whom he tells her that the Turtles have drifted apart. April returns to New York City with a statue for wealthy tycoon Max Winters. Leonardo also returns and April and Casey deliver the statue to Winters. Winters hires Karai and the Foot Clan to search the city for the 13 monsters before the portal opens again. Raphael encounters Casey, who reveals his knowledge of Raphael's double identity and joins him in hunting criminals. Winters, who is actually Yaotl and is still immortal, reanimates his stone generals. Leonardo returns to the sewer lair, meeting Splinter. Splinter forbids the Turtles from fighting until they can act as a team again. While training, the Turtles encounter one of the 13 beasts, Bigfoot, battling the Foot Clan. The Turtles engage Bigfoot, going against Leonardo and Splinter's orders. When Raphael visits Casey, they encounter Vampire Succubor, one of the monsters and witness its capture by the Foot and the Stone Generals, who spot them. Raphael is knocked unconscious. Casey takes Raphael back to the apartment while April calls the Turtles for help and reveals the identities of Yaotl and his generals. After being revived, Raphael suggests they pursue Yaotl, but Leonardo forbids him to go until Splinter gives out the order. Raphael investigates alone.

Leonardo, Donatello, and Michelangelo plan their next move, while Donatello discovers the next portal will open over Winters' tower. Splinter tells Leonardo that his team is incomplete, and that he knows what he must do. After 11 monsters have been captured, General Aguila questions Yaotl's actions. The generals conspire to betray Yaotl, wanting to remain immortal. Raphael encounters the Jersey Devil, one of the remaining monsters but drives it off. Leonardo finds Raphael in his Nightwatcher disguise and, not recognizing him, tries to convince him to stop his vigilante actions. Raphael, still angry, attacks Leonardo, but Leonardo bests him and knocks his helmet off. The two turtles chastise each other’s actions until their anger reaches a boiling point, resulting in a second fight. Raphael breaks Leonardo's swords, but flees in shame when his anger nearly drives him to kill Leonardo. The generals seize Leonardo, intending to substitute him for the 13th missing beast, and Raphael decides to make amends by rescuing Leonardo. As the portal opens, Yaotl discovers his generals' treachery, while Splinter and the Turtles, accompanied by Casey and April, fight their way through the Foot Clan cordon and breach the tower. Yaotl reveals the truth to the heroes: he wants to be free of his curse of immortality. The generals reveal that they wish to preserve their immortality, but also to use the portal to bring in more monsters to conquer the world.

Having refused to betray Yaotl in exchange for serving the Generals, Karai and the Foot Clan work together with April and Casey to search for the final monster, the Sea Monster, while the Turtles fight the generals. Splinter and Yaotl fight off numerous monsters emerging from the portal. April, Casey, and Karai arrive at the tower with the Sea Monster. The Sea Monster crashes into the Generals, dragging them into the portal before it closes. Karai warns them to enjoy their victory while it lasts, claiming they will soon contend with faces from their past, which the Turtles suspect to be the Shredder. She and the rest of the Foot Clan depart. Yaotl, now mortal, honors the Turtles and Splinter, thanking them for fulfilling his wish before dissipating.

Splinter places Yaotl's helmet among his trophy collection (which includes Shredder's helmet, the TGRI canister and the Time Scepter from the live action trilogy), as well as Raphael's "Nightwatcher" helmet and Michelangelo's "Cowabunga Carl" head. As they return to their roles as the shadowy guardians of New York City, Raphael narrates that the Turtles will always be brothers.

• James Arnold Taylor as Leonardo, the oldest of the turtles
• Nolan North as Raphael, the second oldest of the Turtles who has been operating as the vigilante Nightwatcher
• Mitchell Whitfield as Donatello, the second youngest of the Turtles who has been operating as an IT tech support
• Mikey Kelley as Michelangelo, the youngest of the Turtles who has been operating as birthday entertainer "Cowabunga Carl"
• Chris Evans as Casey Jones, April's boyfriend who engages in crimefighting
• Sarah Michelle Gellar as April O'Neil, a female friend of the Turtles who owns an antique shop
• Mako Iwamatsu as Splinter, a mutant rat who is the Turtles' sensei
• Patrick Stewart as Max Winters, a businessman who turned out to be the immortal warlord Yaotl
• Zhang Ziyi as Karai, a female member of the Foot Clan
• John DiMaggio as Colonel Santino, the leader of a group of South American bandits who has been extorting protection money from the local villagers
• Kevin Michael Richardson as General Aguila, the leader of Yaotl's four generals (his name is Spanish for "eagle")
• Paula Mattioli as General Serpiente, one of Yaotl's four generals (her name is Spanish for "snake")
• Fred Tatasciore as General Gato, one of Yaotl's four generals (his name is Spanish for "cat")
• Frank Welker as General Mono, one of Yaotl's generals (his name is Spanish for "monkey")
• Kevin Smith as a diner cook, an unnamed cook at a diner where Nightwatcher fights the Jersey Devil
• Laurence Fishburne as the narrator

Development

A computer graphics imagery (CGI) Teenage Mutant Ninja Turtles (TMNT) movie was first announced in 2000, with John Woo supposedly at the helm. That movie languished in development hell and Woo ultimately moved on to other projects. TMNT, executive produced by the TMNT co-creator Peter Laird, departs from the previous films' live action style and is the first and only CGI film in the series. Writer/director Kevin Munroe said that he wanted to do total CGI instead of live action and CGI turtles because it would be easier for the audience to "suspend disbelief for such an offbeat story" as there would be no break in the reality between CGI and live action. Producer Tom Gray explained that the decision to depart from the live action series was due to escalating budgets for the three films, and with each film making less than its predecessor, a CGI film became a reality. For example, the first film made $135.2 million on a budget of $13.5 million, and the third made $44 million on a budget of $21 million. Orange Sky Golden Harvest's rights to the franchise had expired, and Gray said the question arose there over a CGI TMNT film in 2004.

Writing

Munroe stated in terms of the story line that ideas were floated as extreme as the Turtles being in space, but eventually it just came back to New York City, and the theme of the family that had fallen apart. When developing the screenplay, Munroe wanted to take on a less lighthearted tone or "less Cowabunga" and place an emphasis on dark elements as shown in the original comics to appeal to the mature audience. "I had a very specific tone because mixing that sort of action and comedy is a very specific thing. Most people were just coming and wanting to make it too funny. I think that version of the movie could do really well, but we wanted to do something where it sort of pushes the envelope a little bit more and says that animation is more than just comedic animals bumping into each other and farting!" Munroe said that in design and in the rendering of the animation, he was after the feel of a comic book. Karai was one of Munroe's favoriite characters from the comics and he "was the one who really pushed for Karai" to appear in the film. TMNT co-creator Peter Laird stated it takes place in its own universe separate from the previous films. However, director Munroe says the film exists in the same continuity as the other films, which was supported by the memento wall at the end of the film.

Animation

Development and pre-production for TMNT began in June 2005 at Imagi's Los Angeles facility and the CGI animation was produced in Hong Kong, followed by post-production in Hollywood. In designing the New York backdrop, art director/concept artist Simon Murton stylized the familiar Manhattan skyline and urban landscapes: "We began with cinematic cues from certain black-and-white films from the 1940s and '50s. I really wanted to push the lighting and the environments to create the look and feel of an alternate reality." The animators that worked on the fight sequences were inspired by Hong Kong action films. Animation director Kim Ooi explains said that because of CGI they were able to "push and stylize beyond the limits of live action." Imagi used Maya with Pixar’s RenderMan for the production pipeline’s back-end.

Casting

Jim Cummings was the only past TMNT actor to appear in this film, where he had previously contributed voice-work in the 1987 Teenage Mutant Ninja Turtles series. The film also features three voice actors for Ratchet in the Ratchet & Clank series, Mikey Kelley and Kevin Michael Richardson from the first game, and James Arnold Taylor from the others, playing Michelangelo, General Aguila, and Leonardo, respectively. TMNT would be Mako Iwamatsu's final acting role. Mako was announced as the voice of Splinter at the San Diego Comic-Con on July 20, 2006. He then died the next day, aged 72. A dedication to Mako appears at the end of the film's credits. Although Iwamatsu is the only actor credited in the role, Greg Baldwin performs a substantial portion of Splinter's dialogue in the finished film; Baldwin had already mimicked Iwamatsu's voice when he took up the late actor's role as Iroh in the concurrently-produced cartoon Avatar: The Last Airbender, and used this precedent to successfully lobby to join the cast of TMNT as Splinter following Iwamatsu's death.